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Abstract We report a CO(J= 3−2) detection of 23 molecular clouds in the extended ultraviolet (XUV) disk of the spiral galaxy M83 with the Atacama Large Millimeter/submillimeter Array. The observed 1 kpc²region is at about 1.24 times the optical radius (R₂₅) of the disk, where CO(J= 2–1) was previously not detected. The detection and nondetection, as well as the level of star formation (SF) activity in the region, can be explained consistently if the clouds have the mass distribution common among Galactic clouds, such as Orion A—with star-forming dense clumps embedded in thick layers of bulk molecular gas, but in a low-metallicity regime where their outer layers are CO-deficient and CO-dark. The cloud and clump masses, estimated from CO(3−2), range from 8.2 × 10²to 2.3 × 10⁴M_⊙and from 2.7 × 10²to 7.5 × 10³M_⊙, respectively. The most massive clouds appear similar to Orion A in star formation activity as well as in mass, as expected if the cloud mass structure is common. The overall low SF activity in the XUV disk could be due to the relative shortage of gas in the molecular phase. The clouds are distributed like chains up to 600 pc (or longer) in length, suggesting that the trigger of cloud formation is on large scales. The common cloud mass structure also justifies the use of high-JCO transitions to trace the total gas mass of clouds, or galaxies, even in the high-zuniverse. This study is the first demonstration that CO(3−2) is an efficient tracer of molecular clouds even in low-metallicity environments. 

The sunflower family, Asteraceae, comprises 10% of all flowering plant species and displays an incredible diversity of form. Asteraceae are clearly monophyletic, yet resolving phylogenetic relationships within the family has proven difficult, hindering our ability to understand its origin and diversification. Recent molecular clock dating has suggested a Cretaceous origin, but the lack of deep sampling of many genes and representative taxa from across the family has impeded the resolution of migration routes and diversifications that led to its global distribution and tremendous diversity. Here we use genomic data from 256 terminals to estimate evolutionary relationships, timing of diversification(s), and biogeographic patterns. Our study places the origin of Asteraceae at ∼83 MYA in the late Cretaceous and reveals that the family underwent a series of explosive radiations during the Eocene which were accompanied by accelerations in diversification rates. The lineages that gave rise to nearly 95% of extant species originated and began diversifying during the middle Eocene, coincident with the ensuing marked cooling during this period. Phylogenetic and biogeographic analyses support a South American origin of the family with subsequent dispersals into North America and then to Asia and Africa, later followed by multiple worldwide dispersals in many directions. The rapid mid-Eocene diversification is aligned with the biogeographic range shift to Africa where many of the modern-day tribes appear to have originated. Our robust phylogeny provides a framework for future studies aimed at understanding the role of the macroevolutionary patterns and processes that generated the enormous species diversity of Asteraceae. 

Abstract Asteraceae account for 10% of all flowering plant species, and 35%–40% of these are in five closely related tribes that total over 10 000 species. These tribes include Anthemideae, Astereae, Calenduleae, Gnaphalieae, and Senecioneae, which form one of two enormous clades within Subfamily Asteroideae. We took a phylogenomics approach to resolve evolutionary relationships among these five tribes. We sampled the nuclear and plastid genomes via HybSeq target enrichment and genome skimming, and recovered 74 plastid genes and nearly 1000 nuclear loci, known as Conserved Orthologous Sequences. We tested for conflicting support in both data sets and used network analyses to assess patterns of reticulation to explain the early evolutionary history of this lineage, which has experienced whole‐genome duplications and rapid radiations. We found concordance and conflicting support in both data sets and documented four ancient hybridization events. Due to the timing of the early radiation of this five‐tribe lineage, shortly before the Eocene–Oligocene extinction event (34 MYA), early lineages were likely lost, obscuring some details of their early evolutionary history.